- Use formats that advertisers use the most: 728x90 , 300x250 and 160x600. Will get you good diversity of ads coming in.
- The larger the ad unit the better (as long as it doesn't kill the user experience)
- This might sound "greedy" but if your page is long enough, don't be afraid to use the maximum allowed units on a page (3 adsense for content, 2 link units).
- Merge the ad units with your content wherever applicable. I see a lot of webmasters just putting the ad unit in a "dedicated" location and will not go any further. Instead, use them on natural breaks and divisions in your page.
- I haven't used this as much but place units in the shape of an "F" on the page (if the language is English). Research shows visual hot spots lie in this pattern.
- Make use of ad units close to the navigation, above/below video embeds and in forum posts.
- Go for contrast on dark backgrounds on your ad units; go for colors that blend with lighter backgrounds.
- Analyze hot click areas on your pages and place a single ad in the proximity. Don't stack ads one top of another, that might just hurt conversions.
- Very important: Whenever possible, keep changing ad unit locations every month. Loyal users are very good at filtering out ad units after a while and you need to keep mixing it up by changing units around and experimenting with colors/borders.
Having said that, do listen to your customers. If they start leaving your site because of obtrusive and irrelevant ads, cut down.
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